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syntax - Java: What does ~ mean

In this Java source code I have this line:

if ((modifiers & ~KeyEvent.SHIFT_MASK) != 0) ....

What does the tilde ~ mean?

question from:https://stackoverflow.com/questions/65887146/what-does-the-bitwise-operator-do-exactly-do-in-java

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The Tilde (~) performs a bitwise complement of a numerical value in Java.

See: Bitwise complement (~): inverts ones and zeroes in a number


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